Swans are birds of the genus Cygnus within the family
Anatidae.[4] The swans' closest relatives include the
geese and
ducks. Swans are grouped with the closely related geese in the
subfamilyAnserinae where they form the
tribeCygnini. Sometimes, they are considered a distinct subfamily, Cygninae.
There are six living and many
extinct species of swan; in addition, there is a species known as the
coscoroba swan which is no longer considered one of the true swans. Swans usually
mate for life, although separation sometimes occurs, particularly following nesting failure, and if a mate dies, the remaining swan will take up with another. The number of
eggs in each
clutch ranges from three to eight.[5]
Young swans are known as cygnets, or, less commonly, as swanlings.[10][11][12][13][14][15] The former derives from
Old Frenchcigne or cisne (diminutive suffix et'little'), from the
Latin word cygnus, a variant form of cycnus'swan', itself from the
Greekκύκνοςkýknos, a word of the same meaning.[16][17][18] An adult male is a cob, from
Middle Englishcobbe (leader of a group); an adult female is a pen.[19] A group of swans is called a bevy[citation needed] or a wedge.[20]
Description
Swans are the largest extant members of the waterfowl family
Anatidae and are among the largest flying birds. The largest living species, including the
mute swan,
trumpeter swan, and
whooper swan, can reach a length of over 1.5 m (59 in) and weigh over 15 kg (33 lb). Their wingspans can be over 3.1 m (10 ft).[21] Compared to the closely related geese, they are much larger and have proportionally larger feet and necks.[22] Adults also have a patch of unfeathered skin between the eyes and bill. The sexes are alike in
plumage, but males are generally bigger and heavier than females.[19] The biggest species of swan ever was the extinct Cygnus falconeri, a flightless giant swan known from fossils found on the Mediterranean islands of
Malta and
Sicily. Its disappearance is thought to have resulted from extreme climate fluctuations or the arrival of superior predators and competitors.[23]
The Northern Hemisphere species of swan have pure white plumage, but the Southern Hemisphere species are mixed black and white. The Australian
black swan (Cygnus atratus) is completely black except for the white flight feathers on its wings; the chicks of black swans are light grey. The South American
black-necked swan has a white body with a black neck.[24]
The legs of most swans are typically a dark blackish-grey colour, except for the South American black-necked swan, which has pink legs. Bill colour varies: the four
subarctic species have black bills with varying amounts of yellow, and all the others are patterned red and black. Although birds do not have teeth, swans, like other Anatidae, have beaks with serrated edges that look like small jagged "teeth" as part of their beaks used for catching and eating aquatic plants and algae, but also molluscs, small fish, frogs, and worms.[25] In the mute swan and black-necked swan, both sexes have a fleshy lump at the base of their bills on the upper
mandible, known as the knob, which is larger in males and is condition dependent, changing seasonally.[26][27]
Swans are generally found in temperate environments, rarely occurring in the tropics. Four (or five) species occur in the
Northern Hemisphere, one species is found in Australia, one
extinct species was found in New Zealand and the
Chatham Islands, and one species is distributed in southern South America. They are absent from tropical Asia, Central America, northern South America and the entirety of Africa. One species, the mute swan, has been
introduced to North America, Australia and New Zealand.[22]
Several species are
migratory, either wholly or partly so. The mute swan is a partial migrant, being resident over areas of Western Europe but wholly migratory in Eastern Europe and Asia. The whooper swan and tundra swan are wholly migratory, and the trumpeter swans are almost entirely migratory.[22] There is some evidence that the black-necked swan is migratory over part of its range, but detailed studies have not established whether these movements are long or short-range migration.[28]
Behaviour
Swans feed in water and on land. They are almost entirely herbivorous, although they may eat small amounts of aquatic animals. In the water, food is obtained by up-ending or dabbling, and their diet is composed of the roots, tubers, stems and leaves of aquatic and submerged plants.[22]
A familiar behaviour of swans is that they mate for life, and typically bond even before they reach sexual maturity. Trumpeter swans, for example, can live as long as 24 years and only start breeding at the age of 4–7, forming monogamous
pair bonds as early as 20 months.[29] "Divorce", though rare, does occur; one study of mute swans shows a 3% rate for pairs that breed successfully and 9% for pairs that do not.[30] The pair bonds are maintained year-round, even in gregarious and migratory species like the
tundra swan, which congregate in large flocks in the wintering grounds.[31]
Swans' nests are on the ground near water and about a metre (3') across. Unlike many other ducks and geese, the male helps with the nest construction, and will also take turns incubating the eggs.[32] Alongside the
whistling ducks, swans are the only anatids that will do this. The average egg size (for the mute swan) is 113 × 74 mm (4+1⁄2 x 3 in), weighing 340 g (12 oz), in a clutch size of 4 to 7, and an incubation period of 34–45 days.[33] Swans are highly protective of their nests. They will viciously attack anything that they perceive as a threat to their chicks, including humans. One man was suspected to have drowned in such an attack.[34][35] Swans' intraspecific aggressive behaviour is shown more frequent than interspecific behaviour for food and shelter. The aggression with other species is shown more in
tundra swans.[36]
Systematics and evolution
Evidence suggests that the genus Cygnus evolved in Europe or western Eurasia during the
Miocene, spreading all over the Northern Hemisphere until the
Pliocene. When the southern species branched off is not known. The mute swan is closest to the Southern Hemisphere Cygnus;[37] its habits of carrying the neck curved (not straight) and the wings fluffed (not flush) as well as its bill colour and knob indicate that its closest living relative is the black swan. Given the
biogeography and appearance of the
subgenusOlor, it seems likely that these are of a more recent origin, as evidenced shows by their modern ranges (which were mostly uninhabitable during the
last ice age) and great similarity between the taxa.[1]
Eurasian species that occurs at lower latitudes than the whooper swan and Bewick's swan across Europe into southern Russia, China and the Russian Maritimes. Recent fossil records, according to the
British Ornithologists' Union, show Cygnus olor is among the oldest bird species still extant and it has been upgraded to "native" status in several European countries since this bird has been found in fossil and bog specimens dating back thousands of years. Common temperate Eurasian birds, often semi-domesticated descendants of domestic flocks, are naturalised in the United States and elsewhere.
Europe into southern Russia, China and the Russian Maritimes; introduced populations in North America, Australasia and southern Africa
The largest North American swan. Very similar to the whooper swan (and sometimes treated as a subspecies of it), it was hunted almost to
extinction but has since recovered.
Breeds on the Arctic tundra and winters in more temperate regions of
Eurasia and North America. It consists of two forms, generally considered to be subspecies.
Bewick's swan, Cygnus (columbianus) bewickii is the Eurasian form that migrates from Arctic Russia to western Europe and eastern Asia (China, Korea, Japan) in winter.
Whistling swan, Cygnus (columbianus) columbianus is the North American form.
North America, Eurasia
The
coscoroba swan (Coscoroba coscoroba) from South America, the only species in its genus, is not a true swan. Its phylogenetic position is not fully resolved; it is in some aspects more similar to
geese and
shelducks.[39]
Fossil record
The
fossil record of the genus Cygnus is quite impressive, although allocation to the subgenera is often tentative; as indicated above, at least the early forms probably belong to the C. olor – Southern Hemisphere lineage, whereas the Pleistocene taxa from North America would be placed in Olor. Several prehistoric species have been described, mostly from the Northern Hemisphere. In the Mediterranean, the leg bones of the giant swan (C. falconeri) were found on the islands of
Malta and
Sicily; it may have been over 2 metres from tail to bill, which was taller (though not heavier) than the contemporary local dwarf elephants (Palaeoloxodon falconeri).
Subgenus Chenopis
†
New Zealand swan, Cygnus sumnerensis, an extinct species related to the black swan of Australia
The supposed fossil swans "Cygnus" bilinicus and "Cygnus" herrenthalsi were, respectively, a
stork and some large bird of unknown affinity (due to the bad state of preservation of the referred material).
Many of the cultural aspects refer to the mute swan of Europe. Perhaps the best-known story about a swan is the fairy tale "
The Ugly Duckling". Swans are often a symbol of love or fidelity because of their long-lasting, apparently monogamous relationships. See
Wagner's famous swan-related operas Lohengrin[43] and Parsifal.[44]
As food
Swan meat was regarded as a luxury food in England during the reign of
Elizabeth I. A recipe for baked swan survives from that time: "To bake a Swan Scald it and take out the bones, and parboil it, then season it very well with Pepper, Salt and Ginger, then lard it, and put it in a deep Coffin of Rye Paste with store of Butter, close it and bake it very well, and when it is baked, fill up the Vent-hole with melted Butter, and so keep it; serve it in as you do the Beef-Pie."[45] Swans being raised for food were sometimes kept in
swan pits.
Based on a mistaken belief that the British monarch owns all the swans in Britain, it is popularly believed the British monarch is the only person allowed to eat swans in the United Kingdom.[46][47]
The flag of the
Swiss municipality of
Horgen. The swan symbolizes the town's location at
Lake Zurich and Horgen's political status as administrative capital of
Horgen District.
Other references in classical literature include the belief that, upon death, the mute swan would sing beautifully—hence the phrase
swan song.[49]
The mute swan is also one of the sacred birds of
Apollo, whose associations stem both from the nature of the bird as a symbol of light, as well as the notion of a "swan song". The god is often depicted riding a chariot pulled by or composed of swans in his ascension from
Delos.
In the second century, the Roman poet
Juvenal made a sarcastic reference to a good woman being a "rare bird, as rare on earth as a black swan" (black swans being completely unknown in the Northern Hemisphere until Dutch explorers reached Australia in the 1600s), from which comes the Latin phrase rara avis (rare bird).[50]
Irish lore and poetry
The Irish legend of the
Children of Lir is about a stepmother who transformed her children into swans for 900 years.[51]
In the legend The Wooing of Etain the king of the
Sidhe (subterranean-dwelling, supernatural beings) transforms himself and the most beautiful woman in Ireland, Etain, into swans to escape from the king of Ireland and Ireland's armies. The swan has recently been depicted on an
Irish commemorative coin.
Swans are also present in Irish literature in the poetry of
W. B. Yeats.
"The Wild Swans at Coole" has a heavy focus on the mesmerising characteristics of the swan. Yeats also recounts the myth of Leda and the Swan in
the poem of the same name.
Nordic lore
In
Norse mythology, two swans drink from the sacred
Well of Urd in the realm of
Asgard, home of the
gods. According to the
Prose Edda, the water of this well is so pure and holy that all things that touch it turn white, including this original pair of swans and all others descended from them. The poem Volundarkvida, or the Lay of Volund, part of the
Poetic Edda, also features swan maidens.
In the Finnish epic Kalevala, a swan lives in the Tuoni River located in
Tuonela, the underworld realm of the dead. According to the story, whoever killed a swan would perish as well.
Jean Sibelius composed the Lemminkäinen Suite based on the Kalevala, with the second piece entitled Swan of Tuonela(Tuonelan joutsen). Today, five flying swans are the symbol of the
Nordic countries; the whooper swan (Cygnus cygnus) is the national bird of Finland;[52] and the mute swan is the national bird of Denmark.[53]
Swan Lake ballet
The ballet Swan Lake is among the most canonic of classical ballets. Based on the 1875–76 score by
Pyotr Ilyich Tchaikovsky, the most promulgated choreographic version was created by
Marius Petipa and
Lev Ivanov (1895), the premiere of which was danced by the Imperial Ballet at the
Mariinsky Theater in St. Petersburg. The ballet's lead dual roles of Odette (white swan)/Odile (black swan) represent good and evil[54] and are among the most challenging roles[55] created in Romantic classical ballet. The ballet is in the repertories[56] of ballet companies around the world.
Christianity
A swan is one of the attributes of St.
Hugh of Lincoln, based on the story of a swan who was devoted to him.[57]
Spanish language literature
In
Latin American literature, the Nicaraguan poet
Rubén Darío (1867–1916) consecrated the swan as a symbol of artistic inspiration by drawing attention to the constancy of swan imagery in
Western culture, beginning with the rape of Leda and ending with
Wagner's Lohengrin. Darío's most famous poem in this regard is Blasón – "Coat of Arms" (1896), and his use of the swan made it a symbol for the
Modernismo poetic movement that dominated Spanish language poetry from the 1880s until the First World War. Such was the dominance of Modernismo in Spanish language poetry that the Mexican poet
Enrique González Martínez attempted to announce the end of Modernismo with a
sonnet provocatively entitled, Tuércele el cuello al cisne – "Wring the Swan's Neck" (1910).
Swans are revered in
Hinduism and are compared to saintly persons whose chief characteristic is to be in the world without getting attached to it, just as a swan's feather does not get wet although it is in water. The
Sanskrit word for swan is hamsa and the "Raja Hamsam" or the Royal Swan is the vehicle of Devi
Saraswati, which symbolises the Sattva Guna or purity par excellence. The swan, if offered a mixture of milk and water, is said to be able to drink the milk alone. Therefore, Saraswati, the goddess of knowledge, is seen riding the swan because the swan thus symbolizes Viveka, i.e. prudence and discrimination between the good and the bad or between the eternal and the transient. This is seen as a great quality, as shown by this
Sanskrit verse:
haṁsaḥ śveto bakaḥ śvetaḥ ko bhedo bakahaṁsayoḥ ।
kṣīranīraviveke tu haṁso haṁsaḥ bako bakaḥ ॥
The swan is white, the crane is white, what is the difference between the swan and the crane?
During discriminating between water and milk, the swan is a swan while the crane is a crane!
It is mentioned several times in the
Vedic literature, and persons who have attained great spiritual capabilities are sometimes called
Paramahamsa ("Supreme Swan") on account of their
spiritual grace and ability to travel between various spiritual worlds. In the Vedas, swans are said to reside in the summer on
Lake Manasarovar and migrate to Indian lakes for the winter. They are believed to possess some powers, such as the ability to eat pearls.
Indo-European religions
Swans are intimately associated with the
divine twins in Indo-European religions, and it is thought that in
Proto-Indo-European times, swans were a solar symbol associated with the divine twins and the original Indo-European
sun goddess.[58]
Royal Swans (swans given by Queen Elizabeth II to the city of
Ottawa in 1967, and their progeny)
References
^
abNorthcote, E. M. (1981). "Size difference between limb bones of recent and subfossil Mute Swans (Cygnus olor)". J. Archaeol. Sci. 8 (1): 89–98.
doi:
10.1016/0305-4403(81)90014-5.
^Mallory, J. P.; Adams, D. Q. (2006). The Oxford Introduction to Proto-Indo-European and the Proto-Indo-European World. New York: Oxford University Press. p. 360.
ISBN0-19-928791-0.
^Pauline Somers, Dreams within Dreams, 2004: "Suddenly I feel myself become small as a swanling, tucked into the swan's feathers."
^Hubert Schuurman, People of the Swan, 2007: "Slim ran over to the pond and unleashed the excited swanlings. They followed Moose and Slim to a quiet backwash of the river and glided gracefully into the water. Moose sat down on a little hillock and listened to Slim talking to the swanlings."
^Jack Zipes, The Enchanted Screen, 2011: "The narrative concerns a female duck who hatches four dark ducklings, and when the fifth turns out to be a white swanling, the father duck more or less accuses his wife of adultery."
^Gail Steinberg and Beth Hall, Inside Transracial Adoption, 2011: "Bruno Bettelheim: What this is really about is how you and your own dear swanling can rest securely in the feeling of belonging together when you experience daily life differently because others treat you differently."
^Betina Lindsay, Swan Bride: The Swan Maiden Trilogy — Book One, 2015: "So close, swanling, do not hesitate. Her eyes went to the sun hanging between night and day and then to Wulfsun. I cannot leave him to such a fate. Without me, he is a dead man."
^Horrocks, N., Perrins, C. and Charmantier, A., 2009. Seasonal changes in male and female bill knob size in the mute swan Cygnus olor. Journal of avian biology, 40(5), pp.511-519.
^Schlatter, Roberto; Navarro, Rene A.; Corti, Paulo (2002). "Effects of El Nino Southern Oscillation on Numbers of Black-Necked Swans at Rio Cruces Sanctuary, Chile". Waterbirds. 25 (Special Publication 1): 114–122.
JSTOR1522341.
^Scott, D. K. (1980). "Functional aspects of the pair bond in winter in Bewick's swans (Cygnus columbianus bewickii)". Behavioral Ecology and Sociobiology. 7 (4): 323–327.
doi:
10.1007/BF00300673.
S2CID32804332.
^Scott, Dafila (1995). Swans. Grantown-on-Spey, Scotland: Colin Baxter Photography. p. 51.
ISBN978-0-948661-63-1.
^Boev, Z. 2000. "Cygnus verae sp. n. (Anseriformes: Anatidae) from the Early Pliocene of Sofia (Bulgaria)". Acta zoologica cracovienzia, Krakow, 43 (1–2): 185–192.
^Louchart, Antoine; Vignaud, Patrick; Likius, Andossa; Mackaye, Hassane T.; Brunet, Michel (27 June 2005). "A New Swan (Aves: Anatidae) in Africa, from the Latest Miocene of Chad and Libya". Journal of Vertebrate Paleontology. 25 (2): 384–392.
doi:
10.1671/0272-4634(2005)025[0384:ANSAAI]2.0.CO;2.
JSTOR4524452.
S2CID85860957.
^The ballet Swan Lake is among the most canonic of classical ballets. Based on the 1875-76 score by Pyotr Ilyich Tchaikovsky, the most promulgated choreographic version was created by Marius Petipa and Lev Ivanov (1895), the premiere of which was danced by the Imperial Ballet at the Mariinsky Theater in St. Petersburg. The ballet's lead dual roles of Odette/Odile represent good and evil, and are among the most challenging roles created in Romantic classical ballet.
Louchart, Antoine; Mourer-Chauviré, Cécile; Guleç, Erksin; Howell, Francis Clark & White, Tim D. (1998): L'avifaune de Dursunlu, Turquie, Pléistocène inférieur: climat, environnement et biogéographie. C. R. Acad. Sci. Paris IIA327(5): 341–346. [French with English abridged version]
doi:
10.1016/S1251-8050(98)80053-0