Many non-avian
dinosaurs were feathered. Direct evidence of feathers exists for the following species, listed in the order currently accepted evidence was first published. In all examples, the evidence described consists of feather impressions, except those genera inferred to have had feathers based on skeletal or chemical evidence, such as the presence of quill knobs (the anchor points for wing feathers on the forelimb) or a
pygostyle (the fused vertebrae at the tail tip which often supports large feathers).[1]
Note that the filamentous structures in some
ornithischian dinosaurs (Psittacosaurus, Tianyulong and Kulindadromeus) and the pycnofibres found in some
pterosaurs may or may not be
homologous with the feathers of theropods.[39][74]
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^Czerkas, S.A., and Yuan, C. (2002). "An arboreal maniraptoran from northeast China." Pp. 63-95 in Czerkas, S.J. (Ed.), Feathered Dinosaurs and the Origin of Flight. The Dinosaur Museum Journal 1. The Dinosaur Museum, Blanding, U.S.A.
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^Ji S.; Ji Q.; Lu J.; Yuan C. (2007). "A new giant compsognathid dinosaur with long filamentous integuments from Lower Cretaceous of Northeastern China". Acta Geologica Sinica. 81 (1): 8–15.
^He T.; Wang X.-L.; Zhou Z.-H. (2008). "A new genus and species of caudipterid dinosaur from the Lower Cretaceous Jiufotang Formation of western Liaoning, China". Vertebrata PalAsiatica. 46 (3): 178–189.
^Xu, Xing; Zhao, Qi; Norell, Mark; Sullivan, Corwin; Hone, David; Erickson, Gregory; Wang, XiaoLin; Han, FengLu; Guo, Yu (2008). "A new feathered maniraptoran dinosaur fossil that fills a morphological gap in avian origin". Chinese Science Bulletin. 54 (3): 430–435.
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^Xu, Xing (2020), "Filamentous Integuments in Nonavialan Theropods and Their Kin: Advances and Future Perspectives for Understanding the Evolution of Feathers", in Foth, Christian; Rauhut, Oliver W. M. (eds.), The Evolution of Feathers, Fascinating Life Sciences, Springer International Publishing, pp. 67–78,
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^Xing Xu; Hailu You; Kai Du; Fenglu Han (28 July 2011). "An Archaeopteryx-like theropod from China and the origin of Avialae". Nature. 475 (7357): 465–470.
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^Xu, Xing; Wang, Kebai; Zhang, Ke; Ma, Qingyu; Xing, Lida; Sullivan, Corwin; Hu, Dongyu; Cheng, Shuqing; Wang, Shuo (2012). "A gigantic feathered dinosaur from the Lower Cretaceous of China". Nature. 484 (7392): 92–95.
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^Darla K. Zelenitsky; François Therrien; Gregory M. Erickson; Christopher L. DeBuhr; Yoshitsugu Kobayashi; David A. Eberth; Frank Hadfield (2012). "Feathered Non-Avian Dinosaurs from North America Provide Insight into Wing Origins". Science. 338 (6106): 510–514.
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^Ji Qiang; Lü Jun-Chang; Wei Xue-Fang; Wang Xu-Ri (2012). "A new oviraptorosaur from the Yixian Formation of Jianchang, Western Liaoning Province, China". Regional Geology of China (12): 2102–2107.
^Pascal Godefroit; Andrea Cau; Hu Dong-Yu; François Escuillié; Wu Wenhao; Gareth Dyke (2013). "A Jurassic avialan dinosaur from China resolves the early phylogenetic history of birds". Nature. 498 (7454): 359–362.
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^Xu, X.; Zheng, X.; Sullivan, C.; Wang, X.; Xing, L.; Wang, Y.; Zhang, X.; O'Connor, J. K.; Zhang, F.; Pan, Y. (2015). "A bizarre Jurassic maniraptoran theropod with preserved evidence of membranous wings". Nature. 521 (7550): 70–3.
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^Van Der Reest, Aaron J.; Wolfe, Alexander P.; Currie, Philip J. (2016). "A densely feathered ornithomimid (Dinosauria: Theropoda) from the Upper Cretaceous Dinosaur Park Formation, Alberta, Canada". Cretaceous Research. 58: 108–117.
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^Robert A. DePalma; David A. Burnham; Larry D. Martin; Peter L. Larson; Robert T. Bakker (2015). "The first giant raptor (Theropoda: Dromaeosauridae) from the Hell Creek Formation". Paleontological Contributions. 14: 1–15.
^Ulysse Lefèvre; Andrea Cau; Aude Cincotta; Dongyu Hu; Anusuya Chinsamy; François Escuillié; Pascal Godefroit (2017). "A new Jurassic theropod from China documents a transitional step in the macrostructure of feathers". The Science of Nature. 104 (9–10): Article number 74.
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