Genus of gastropods
Clavus is a
genus of small
sea snails, marine
gastropod
mollusks in the
family
Drilliidae.
[1]
[2]
Description
Apart from the general characteristics of the genera in the family
Drilliidae, the species in the genus Clavus have a shell with peripheral tubercles, spines or wing-like processes.
[3] The
aperture is rather large. The
outer lip is produced below the sinus.
[4]
The shell of Clavus is characterised by the following features:— Flat indefinite fasciole, indicated only by the curve of growth lines. A smooth subulate
protoconch. An insinuation of the outer lip, near the base, like that of
Strombus. A major
sculpture of prominent arched scales on the shoulder, and a minor sculpture of dense, microscopic, waved, spiral striae.
[5]
Most species in this genus have a dorsal
varix. But this is absent in a few species such as
Clavus beckii,
Clavus humilis and
Clavus pica.
G.W. Tryon correctly preserved the genus for smooth specimens with a short
body whorl, long spire, nodulous shoulder, no spiral sculpture, a wide, deep
anal sulcus adjacent to the suture and, in the completely adult, a marked subsutural callus on the body.
[6]
Distribution
The species in this genus occur in the
Red Sea, In the Indian Ocean off
Mozambique; also off
Indonesia,
New Caledonia,
Papua New Guinea, the
Philippines,
Solomon Islands,
Japan and Australia (
New South Wales,
Northern Territory,
Queensland,
Western Australia).
Species
According to the
World Register of Marine Species (WoRMS) the following species with valid names are included within the genus Clavus :
[7]
-
Clavus aenigmaticus Wells, 1991
-
Clavus aglaia (Dall, 1918)
-
Clavus alboroseus Horro, Gori, Rosado & Rolán, 2021
-
Clavus albotuberculatus (Schepman, 1889)
-
Clavus andreolbrichi Fedosov & Puillandre, 2020
-
Clavus androyensis (Bozzetti, 2007)
-
Clavus angulatus Stahlschmidt, Poppe & Tagaro, 2018
-
Clavus basipunctatus (Kilburn, 1988)
-
Clavus beckii (Reeve, 1842)
-
Clavus berenice (Dall, 1918) (synonym: Clavus spinosa Smith, E.A., 1882 )
-
Clavus biancae (Bozzetti, 2008)
-
Clavus bilineatus (Reeve, 1845)
-
Clavus boucheti Kilburn, Fedosov & Kantor, 2014
-
Clavus brianmayi Fedosov & Puillandre, 2020
-
Clavus burnupi (G. B. Sowerby III, 1897)
-
Clavus cadenasi (Clench & Aguayo, 1939)
-
Clavus canalicularis (Röding, 1798)
-
Clavus candens (Smith E. A., 1879)
-
Clavus cantharis (Reeve, 1845)
-
Clavus clara (Reeve, 1845)
-
Clavus clavata (Sowerby II, 1870)
-
Clavus cloveri (Poppe, Tagaro & Goto, 2018)
-
Clavus coffea (Smith E. A., 1882)
-
Clavus cygneus (Melvill & Standen, 1897)
-
Clavus davidgilmouri Fedosov & Puillandre, 2020
-
Clavus delphineae Kilburn, Fedosov & Kantor, 2014
-
Clavus devexistriatus Kilburn, Fedosov & Kantor, 2014
-
Clavus dolichurus Stahlschmidt, Poppe & Tagaro, 2018
-
Clavus dubrucqueae T. Cossignani, 2021
-
Clavus ebur (Reeve, 1845)
-
Clavus exasperatus (Reeve, 1843)
-
Clavus exilis (Pease, 1868)
-
Clavus falcicosta (Barnard, 1958)
-
Clavus flammulatus Montfort, 1810
-
Clavus formosus (Reeve, 1846)
-
Clavus fulvus (Hinds, 1843)
-
Clavus fusconitens (Sowerby III, 1901)
-
Clavus glaucozona Kilburn & Dekker, 2008
-
Clavus groschi Kilburn, 1988
-
Clavus herberti (Kilburn, 1988)
-
Clavus heryi (Bozzetti, 2007)
-
Clavus hewittae Wells, 1991
-
Clavus hottentotus (E. A. Smith, 1882)
-
Clavus humilis (E. A. Smith, 1879)
-
Clavus hylikos Kilburn, Fedosov & Kantor, 2014
-
Clavus idae (Poppe, Tagaro & Goto, 2018)
-
Clavus infrafusca (G.B. Sowerby III, 1893)
-
Clavus infuscatus Kilburn & Dekker, 2008
-
Clavus isibopho (Kilburn, 1988)
-
Clavus isowai Poppe, Tagaro & Goto, 2018
-
Clavus japonicus (Lischke, 1869)
-
Clavus johnsoni (Bartsch, 1934)
-
Clavus juani Horro, Gori, Rosado & Rolán, 2021
-
Clavus kirkhammetti Fedosov & Puillandre, 2020
-
Clavus laetus (Hinds, 1843)
-
Clavus lamberti (Montrouzier, 1860)
-
Clavus maestratii Kilburn, Fedosov & Kantor, 2014
-
Clavus malva (Morassi, 1998)
-
Clavus minutissimus Stahlschmidt, Poppe & Tagaro, 2018
-
Clavus moquinianus (Montrouzier, 1874)
-
Clavus nodifera (Pease, 1860)
-
Clavus nodulosa (Pease, 1863)
-
Clavus obliquatus (Reeve, 1845)
-
Clavus obliquicostatus (Reeve, 1845)
-
Clavus occiduus Wells, 1991
-
Clavus papilio (Kilburn, 1988)
-
Clavus paroeca (Melvill, 1923)
-
Clavus particolor Stahlschmidt, Poppe & Tagaro, 2018
-
Clavus peristera Melvill, 1927
-
Clavus pica (Reeve, 1843)
-
Clavus picoides Kilburn, Fedosov & Kantor, 2014
-
Clavus powelli Kay, 1979
-
Clavus protentus Hervier, 1896
-
Clavus pulicarius Wells, 1991
-
Clavus pusilla (Garrett, 1873)
-
Clavus putillus (Reeve, 1845)
-
Clavus quadratus (Kilburn, 1988)
-
Clavus rissoiniformis Kay, 1979
-
Clavus roseofuscus (Bozzetti, 2007)
-
Clavus rugizonatus Hervier, 1896
-
Clavus sacra (Reeve, L.A., 1845)
-
Clavus similis Stahlschmidt, Poppe & Tagaro, 2018
-
Clavus squamiferus Kilburn, Fedosov & Kantor, 2014
-
Clavus subobliquatus (E. A. Smith, 1879)
-
Clavus subtilifasciatus Horro, Gori, Rosado & Rolán, 2021
-
Clavus suduirauti Bozzetti, 1997
-
Clavus sulekile (Kilburn, 1988)
-
Clavus triantoniorum Horro, Gori, Rosado & Rolán, 2021
-
Clavus unizonalis (Lamarck, 1822)
-
Clavus velcolorum Horro, Gori, Rosado & Rolán, 2021
-
Clavus vibicinus (Helbling, 1779)
-
Clavus virginieae Kilburn, Fedosov & Kantor, 2014
-
Clavus wilmeri (Smith E. A., 1879)
- Species brought into synonymy
- Clavus abdera (Dall, 1919): synonym of
Crassispira abdera (Dall, 1919)
- Clavus aeneus (Hedley, 1922): synonym of
Antiguraleus aeneus (Hedley, 1922)
- Clavus alabaster (Reeve, 1843): synonym of
Inquisitor alabaster (Reeve, 1843)
- Clavus auriculifera Lamarck, J.B.P.A. de, 1816: synonym of
Clavus canalicularis (Röding, 1798)
- Clavus candidulus Hedley, 1922: synonym of
Splendrillia candidulus (Hedley, 1922)
- Clavus costatus Hedley, 1922: synonym of
Graciliclava costata (Hedley, 1922)
- Clavus crassa (E.A. Smith, 1888): synonym of Clavus aglaia (Dall, 1918)
- Clavus enna (Dall, 1918): synonym of
Drillia enna (Dall, 1918)
- Clavus filicinctus (E. A. Smith, 1882): synonym of
Horaiclavus filicinctus (E. A. Smith, 1882)
- Clavus flavidulus (Lamarck, 1822): synonym of
Clathrodrillia flavidula (Lamarck, 1822)
- Clavus flexus Shuto, 1983: synonym of
Plagiostropha flexus (Shuto, 1983)
- Clavus gibberulus Hervier, 1896 : synonym of
Drillia gibberulus (Hervier, 1896)
- Clavus inclinata (Sowerby III, 1893): synonym of
Iredalea inclinata (Sowerby III, 1893)
- Clavus lacertosus (Hedley, 1922): synonym of
Bathytoma lacertosus Hedley, 1922
- Clavus leforestieri Hervier, 1896: synonym of Clavus obliquicostatus (Reeve, 1845)
- Clavus maravignae (Bivona Ant. in Bivona And., 1838): synonym of
Crassopleura maravignae (Bivona Ant. in Bivona And., 1838)
- Clavus mighelsi Kay, 1979: synonym of
Pyrgocythara mighelsi (Kay, 1979)
- Clavus mighelsi of authors: synonym of Clavus humilis (E. A. Smith, 1879)
- Clavus opalus (Reeve, 1845): synonym of
Plagiostropha opalus (Reeve, 1845)
- Clavus pseudoprincipalis (Yokoyama, 1920): synonym of
Inquisitor pseudoprincipalis (Yokoyama, 1920)
- Clavus pulchellus (Reeve, 1845): synonym of
Fenimorea halidorema Schwengel, 1940
- Clavus quintuplex (Melvill, 1927): synonym of
Plagiostropha quintuplex Melvill, 1927
- Clavus regius Habe & Murakami, 1970: synonym of
Drillia regia (Habe & Murakami, 1970)
- Clavus undatus (Hedley, 1907): synonym of
Austroclavus undatus (Hedley, 1907)
- Clavus viduus (Reeve, 1845): synonym of
Clavus unizonalis (Lamarck, 1822)
- Species inquirenda
The Indo-Pacific Molluscan Database adds the following species with names in current use :
- Subgenus Clavus
References
- Montfort, 1810 [Conchyliologie systématique, 2: 434]
- Habe T. (1958). On the radulae of Japanese marine gastropods (4). Venus. 20(1): 43-60, pls 2-3
External links
Wikimedia Commons has media related to
Clavus.